Home / Articles in Palaeobotany / Tertiary Palaeobotany

Ü. Akkemik et al. (2023): Wood and leaf remains of palms with affinities to Sabal Adans., from the middle Eocene of Turkey. Free access, Turkish Journal of Botany, 47: 50-60. https://doi.org/10.55730/1300-008X.2743 Available at: https://journals.tubitak.gov.tr/

! J.M. Anderson et al. (1999): Patterns of Gondwana plant colonisation and diversification. In PDF, Journal of African Earth Sciences, 28: 145-167.
See also here.

A. Andruchow-Colombo et al. (2019): Oldest record of the scale-leaved clade of Podocarpaceae, early Paleocene of Patagonia, Argentina. In PDF, Alcheringa, 43. See also here.

A. Andruchow-Colombo et al. (2019): A South American fossil relative of Phyllocladus: Huncocladus laubenfelsii gen. et sp. nov.(Podocarpaceae), from the early Eocene of Laguna del Hunco, Patagonia, Argentina. Australian Systematic Botany, 32: 290–309. See also here (in PDF).

! Arctic Plant Fossils (hosted by the Institute of Botany, Chinese Academy of Sciences).
This interactive illustrated catalogue of Cretaceous and Paleogene Arctic plant fossils is the outcome of a project from the United States Geological Survey and administered through the University of Oxford, UK, and the Imperial College, London.
Images of the fossils and information on where they were found can be accessed through interactive maps,
or navigating the site by means of tables of taxonomic names (if known), museum collections or the collectors and researchers who worked on them. Excellent!

A.T. Aung et al. (2020): Three new fossil records of Equisetum (Equisetaceae) from the Neogene of south-western China and northern Vietnam. In PDF, PhytoKeys, 138: 3–15.

N.N.A. Bayam et al. (2018): Further contributions to the early Miocene forest vegetation of the Galatian Volcanic Province, Turkey. Free access, Palaeontologia Electronica.

S. Beurel et al (2024): First flower inclusion and fossil evidence of Cryptocarya (Laurales, Lauraceae) from Miocene amber of Zhangpu (China). In PDF, Fossil Record, 27: 1–11.
See likewise here and there.
"... We here described the first Cenozoic Lauraceae flower of Asia and confirmed the presence of Cryptocarya in the Miocene Zhangpu flora
[...] We scanned the specimen using synchrotron radiation-based micro-computed tomography (SRìCT) and then compared the fossil with extant flowers of the genus ..."

H.J.B. Birks and W. Tinner (2016): Past forests of Europe. In PDF, European Atlas of Forest Tree Species.

J. Blanchard et al. (2016): Fruits, seeds and flowers from the Bovay and Bolden clay pits (early Eocene Tallahatta Formation, Claiborne Group), northern Mississippi, USA. In PDF, Palaeontologia Electronica. See also here.

J.M. Bouchal et al. (2020): Messinian vegetation and climate of the intermontane Florina–Ptolemais–Servia Basin, NW Greece inferred from palaeobotanical data: how well do plant fossils reflect past environments? Open access, R. Soc. Open Sci. 7: 192067.

Department of Geological Sciences, University of Colorado, Boulder, CO: Web-based instruction. Annotated links to information on using the web to teach. Go to: CzPaleobotany. Go to: Cenozoic Elevation of the Rocky Mountains, Paleobotanical Methods. About fossil classification (nearest living relative, physiognomy and CLAMP) and climate and elevation analysis.
These expired links are now available through the Internet Archive´s Wayback Machine.

M. Brea et al. (2015): Reconstruction of a fossil forest reveals details of the palaeoecology, palaeoenvironments and climatic conditions in the late Oligocene of South America. In PDF, Palaeogeography, Palaeoclimatology, Palaeoecology, 418: 19-42.

G.E. Budd et al. (2021): The use of geological and paleontological evidence in evaluating plant phylogeographic hypotheses in the Northern Hemisphere Tertiary. Free access. See also here (in PDF).

R.J. Burnham and K.R. Johnson (2004): South American palaeobotany and the origins of neotropical rainforests. In PDF, Phil. Trans. R. Soc. Lond., B 359: 1595-1610.

J. Carrión et al. (2022): Palearctic floras and vegetation of the Cenozoic: A tribute to Zlatko Kvacek. Review of Palaeobotany and Palynology, 306.
See also here.

H.Y. Chen et al. (2020): The Oligocene Equisetum from Qaidam Basin, Northeastern Tibetan Plateau in China and its implications. In PDF, Historical Biology, DOI: 10.1080/08912963.2020.1830280.

Y.-S. Chen et al. (2018): Is the East Asian flora ancient or not? In PDF, National Science Review, 0: 1–13. See also here

C.J. Cleal & B.A. Thomas (2001): Introduction to the Mesozoic and Tertiary palaeobotany of Great Britain. PDF file, from: Cleal, C.J., Thomas, B.A., Batten, D.J. & Collinson, M.E., (2001), Mesozoic and Tertiary Palaeobotany of Great Britain, Geological Conservation Review Series, No. 22, Joint Nature Conservation Committee, Peterborough, 335 pages, illustrations, A4 hardback, ISBN 1 86107 489 1.
Still available via Internet Archive Wayback Machine.
Note figure 1.1: The potential process involved in a plant fragment passing into the fossil record.
Figure 1.2 Summary of modes and nomenclature of plant fossil preservation.

C.J. Cleal et al. (2001):
Geological Conservation Review Series (GCR), Joint Nature Conservation Committee (JNCC): Mesozoic and Tertiary Palaeobotany of Great Britain (2001). PDF files, GCR Volume No. 22.
This expired link is now available through the Internet Archive´s Wayback Machine.
In chapter 1 a brief explanation is given of how plant fossils are formed, and how palaeobotanists study and name them.

! M.E. Collinson (2002): The ecology of Cainozoic ferns. In PDF, Review of Palaeobotany and Palynology, 119: 51-68.
See also here.
! Note table 1: Summary of key conclusions concerning the ecology of Cenozoic ferns.

! T. Denk et al, (2023): Cenozoic migration of a desert plant lineage across the North Atlantic. Free access, New Phytologist, 238: 2668–2684.
Note figure 5: Timing and mode of intercontinental Madrean–Tethyan disjunctions of sclerophyllous plants.
"... The fossil record suggests that Vauquelinia, currently endemic to arid and subdesert environments, originated under seasonally arid climates in the Eocene of western North America and subsequently crossed the Paleogene North Atlantic land bridge (NALB) to Europe. This pattern is replicated by other sclerophyllous, dry-adapted and warmth-loving plants ..."
Also worth checking out:
! R.S. Hill and R. Khan (2023): Past climates and plant migration – the significance of the fossil record.
A commentary on Denk et al. (2023).

Thomas Denk et al. (2019): Comment on "Eocene Fagaceae from Patagonia and Gondwanan legacy in Asian rainforests". Free access, Science, 366. DOI: 10.1126/science.aaz2189.
"... extensive paleobotanical records of Antarctica and Australia lack evidence of any Fagaceae, and molecular patterns indicate shared biogeographic histories of Castanopsis, Castanea, Lithocarpus, and Quercus subgenus Cerris, making the southern route unlikely."

! Denver Museum of Nature and Science, Denver, Colorado: DMNS Paleobotany Collection. This website contains over 1000 images of fossil plants spanning the late Cretaceous through early Eocene from the Western Interior of North America. Go to: Identification Flow Chart, or start with Morphotype a Flora. A guide to morphotyping (or binning) a fossil flora step-by-step.

! Digiphyll (State Museum of Natural History Stuttgart).
Digiphyll is designed as an educational portal to provide effective assistance in identifying fossil plant material. Excellent! Please note:
Manual: How to use this portal (in PDF).
Glossary: Leaf morphology (in PDF).
! Worth to check out: The fact sheets (PDF files, 36 taxa). Click the button "Downloads".

R.M. Dillhoff et a. (2013): The Eocene Thomas Ranch flora, Allenby Formation, Princeton, British Columbia, Canada. In PDF, Botany, 91.
See also here.

D. Dimitrov et al. (2023): Diversification of flowering plants in space and time. Free access, Nature Communications, 14.
"... Using a newly generated genus-level phylogeny and global distribution data for 14,244 flowering plant genera, we describe the diversification dynamics of angiosperms through space and time. Our analyses show that diversification rates increased throughout the early Cretaceous and then slightly decreased or remained mostly stable until the end of the Cretaceous–Paleogene mass extinction event 66 million years ago. After that, diversification rates increased again towards the present ..."

W.N. Ding et al. (2018): A new fossil species of Cryptomeria (Cupressaceae) from the Rupelian of the Lühe Basin, Yunnan, East Asia: Implications for palaeobiogeography and palaeoecology. Abstract, Review of Palaeobotany and Palynology, 248: 41-51.
Also of interest in this context:
Pflanzliche Botschaften aus der Urzeit (by Tamara Worzewski, November 08, 2022, Spektrum.de, in German).

M. Dolezych and W. Schneider (2006): Inkohlte Hölzer und Cuticulae dispersae aus dem 2. Miozänen Flözhorizont im Tagebau Welzow (Lausitz)–Taxonomie und vergleichende feinstratigraphisch-fazielle Zuordnung. PDF file, in German. Zeitschrift für geologische Wiss. See also here.
Note photographs of tree stumps on plate 2.

M. Dolezych (2005): Koniferenhölzer im Lausitzer Flöz und ihre ökologische Position. PDF file, thesis, in German, with English summary (starting on PDF page 27). LLP Contributions Series, 19.

M. Eberlein (2015): Bestimmungs- und Verbreitungsatlas der Tertiärflora Sachsens – Angiospermenblätter und Ginkgo. PDF file (in German). Thesis, University of Dresden (in German). First part of a reference book of the Tertiary flora of Saxony.
See also here.
Please take notice:
! Starting on PDF page 30: "Kurzer Abriss der tertiärpaläobotanischen Forschung in Sachsen". The history of saxonian tertiary palaeobotany in brief.

Beth Ellis et al. (2009): Manual of Leaf Architecture. Book announcement. The link is to a version archived by the Internet Archive´s Wayback Machine.
! See also here and there.

B. Erdei et al. (2022): The late Oligocene macroflora of Zsámbék, central Hungary. Open access, Fossil Imprint, 78: 298–309.

B. Erdei et al. (2019): First cycad seedling foliage from the fossil record and inferences for the Cenozoic evolution of cycads. In PDF, Biol. Lett., 15: 20190114. See also here.

B. Erdei and S.R. Manchester (2015): Ctenis clarnoensis sp. n., an Unusual Cycadalean Foliage from the Eocene Clarno Formation, Oregon. In PDF, Int. J. Plant Sci., 176: 1–43.

B. Erdei et al. (2009): The buried Miocene forest at Bükkábrány, Hungary. In PDF, Review of Palaeobotany and Palynology, 155: 69–79.
See also here.
Note plate 1: In situ upright stumps in the opencast mine at Bükkábrány.
"... fifteen ‘in situ’ stumps standing at their original position ..."

Eurekalert.org: Evidence is weak for tropical rainforest 65 million years ago in Africa´s low-latitudes. Results of Bonnie F. Jacobs at Southern Methodist University in Dallas. See also: here (the scarcity of paleobotanical sites in Africa), and there.

Evolving Earth Foundation Issaquah, WA:
The Evolving Earth Foundation is a small Private Operating Foundation with focus on the Tertiary paleobotany of western North America, with interests in systematics, paleoecology and biogeography. Go to:
The Fossil Plant Image Collection.
With line drawings of leaves from the Tertiary.

David K. Ferguson et al. (2009): The taphonomy of a remarkable leaf bed assemblage from the Late Oligocene-Early Miocene Gore Lignite Measures, southern New Zealand. PDF file, International Journal of Coal Geology.
Snapshot taken by the Internet Archive´s Wayback Machine.

Jane E. Francis and Imogen Poole (2002): Cretaceous and early Tertiary climates of Antarctica: evidence from fossil wood. PDF file, Palaeogeography, Palaeoclimatology, Palaeoecology, 182: 47-64.

M. Friedman and G. Carnevale (2018): The Bolca Lagerstätten: shallow marine life in the Eocene. In PDF, Journal of the Geological Society, 175: 569–579.
See likewise here.
"... Famous for its fishes, the localities of Bolca also yield diverse invertebrate faunas and a rich, but relatively understudied flora ..."

M.K. Futey et al. (2012): Arecaceae Fossil Fruits from the Paleocene of Patagonia, Argentina. In PDF, The Botanical Review, 78: 205–234.
See also here.

Maria A. Gandolfo-Nixon, The Liberty Hyde Bailey Hortorium Herbarium (BH) at Cornell University: Patagonian Paleofloras.

N. Gentis et al. (2024): First fossil woods and palm stems from the mid-Paleocene of Myanmar and implications for biogeography and wood anatomy. Open access, Am J Bot., 111. https://doi.org/10.1002/ajb2.16259.

R.S. Hill and R. Khan (2023): Past climates and plant migration – the significance of the fossil record. Free access, New Phytologist.
This article is a Commentary on Denk et al. (2023):
Cenozoic migration of a desert plant lineage across the North Atlantic.

C.T. Gee et al. (2022): First water lily, a leaf of Nymphaea sp., from the Miocene Clarkia flora, northern Idaho, USA: Occurrence, taphonomic observations, floristic implications. In PDF, Fossil Imprint, 78: 288–297.
See likewise here
"... it would be expected that fossil water lily leaves in an ancient pond or lake would make up a sizeable proportion of any fossil plant assemblage
[...] the remains of aquatic macrophytes are strangely uncommon in freshwater deposits of the Cenozoic [...] even if the occurrence of one nymphaealean leaf or seed indicates that water lilies had colonized that body of water.
[...] In other ancient freshwater deposits well known as conservation lagerstätten [...] water lily fossils make up a very small proportion of the entire fossil flora ..."

L. Hernandez-Sandoval et al. (2023): Nichima gen. nov. (Alismataceae) based on reproductive structures from the Oligocene�]Miocene of Mexico. Open access, American Journal of Botany, 110.
Note figure 7: Fossil flower reconstructions and proposed inflorescence organization.
"... Two fossil flowers preserved in amber from the Miocene ..."

G.L. Hoffman (2002): Paleobotany and paleoecology of the Joffre Bridge Roadcut locality (Paleocene), Red Deer, Alberta. 2^nd edition. M.Sc. thesis (in PDF), University of Alberta, Edmonton, Alberta.

! M.M. Howell et al. (2022): A modified, step-by-step procedure for the gentle bleaching of delicate fossil leaf cuticles. Open access, Fossil Imprint, 78: 445–450.
See also here. "... Previously, the fossil conifer needles from Miocene lignites were consistently destroyed by the use of Schulze’s reagent and produced unusable results with only 5–10% sodium hypochlorite solution. By using the modified weak bleach method given here, large areas of cuticles could be prepared, remained intact, and yielded good diagnostic information on the leaves. ..."

Y. Huang et al. (2015): Distribution of Cenozoic plant relicts in China explained by drought in dry season. Open access, Scientific Reports, 5.

P. Jardine (2011): The Paleocene-Eocene Thermal Maximum. In PDF, Palaeontology Online. See also here.

Kenneth G. Karol, Richard M. McCourt, Matthew T. Cimino, and Charles F. Delwiche, Science Magazine: The Closest Living Relatives of Land Plants. This analysis supports the hypothesis that the land plants are placed phylogenetically within the Charophyta, identifies the Charales (stoneworts) as the closest living relatives of plants.

A.A. Klymiuk et al. (2016): Mesozoic and Cenozoic plant evolution and biotic change: Introduction and dedication. In PDF, Botany, 94. See also here and there (table of contents).

V.A. Korasidis et al. (2022): Global Changes in Terrestrial Vegetation and Continental Climate During the Paleocene-Eocene Thermal Maximum. Free access, Paleoceanography and Paleoclimatology, 37: e2021PA004325.
Note figure 1: Global paleogeographic reconstruction for 56 Ma illustrating the positions and general depositional settings of PETM palynofloral sites.

J. Kovar-Eder and V. Teodoridis (2018): The Middle Miocene Central European plant record revisited; widespread subhumid sclerophyllous forests indicated. In PDF, Fossil Imprint, 74: 115–134.

K.A. Krakhmal (2023): Paleobiological Methods for Studying the History of Early Anthropogenes in Fergana. Free access. Texas Journal of Philology, Culture and History, 15.

S. Kundu et al. (2023): Evidence of the oldest extant vascular plant (horsetails) from the Indian Cenozoic. Free access, Plant Diversity.

! E. Kustatscher et al. (2022): A whole-plant specimen of the marine macroalga Pterigophycos from the Eocene of Bolca (Veneto, N-Italy). Open access, Fossil Imprint, 78: 145–156.
Note text-figure 5: Reconstruction drawing of Pterigophycos sp. thallus growing on a rock surface.

S.A. Little et al. (2004): Duabanga-like leaves from the Middle Eocene Princeton chert and comparative leaf histology of Lythraceae sensu lato. Open access, American Journal of Botany, 91: 1126-1139.

M.I. Lönartz et al. (2023): Palaeoenvironmental conditions for the natural vulcanization of the Eocene “monkeyhair” laticifers from Geiseltal, Germany, as elucidated by Raman spectroscopy. Free access, Palaeobiodiversity and Palaeoenvironments, 103: 681–693.
"... Raman spectra of the fossil laticifers are virtually identical to that of rubber (cis-1,4-polyisoprene) with additional bands demonstrating sulfur vulcanization. Raman spectra from the surrounding lignite and existing Raman-based carbon thermometers, currently calibrated down to about 100 °C, clearly indicate that these samples were never exposed to temperatures higher than the surrounding lignite. These results directly validate the previous hypothesis of fossilization through natural vulcanization ..."

! S. Magallón (2009): Flowering plants (Magnoliophyta). PDF file, In: S.B. Hedges and S. Kumar (eds.): The Timetree of Life (see here).

D.H. Mai (2007): The floral change in the Tertiary of the Rhön mountains (Germany). In PDF, Acta Palaeobotanica, 47: 135–143.

E. Martinetto and L. Macaluso (2018): Quantitative application of the Whole-Plant Concept to the Messinian – Piacenzian flora of Italy. In PDF, Fossil Imprint, 74: 77–100.

R.W. Mathewes et al. (2023): Plant Megafossils, Palynomorphs, and Paleoenvironment from the Late Middle to Late Eocene Burnaby Mountain Flora, Huntingdon Formation, British Columbia, Canada. Free access, International Journal of Plant Sciences, 184: 157-235. See also:
SFU paleobotanist discovers ancient fossils on Burnaby Mountain (The Peak, 2023).
B.C. paleobotanist rediscovers Burnaby plant fossils stored at SFU, writes paper (by Elizabeth McSheffrey, Global News, April 14, 2023).

J. McCoy et al. (2022): Middle Miocene (Serravallian) wetland development on the northwest edge of Europe based on palynological analysis of the uppermost Brassington Formation of Derbyshire, United Kingdom. In PDF, Palaeogeography, Palaeoclimatology,Palaeoecology, 603.
See also here.

! V. Mosbrugger et al. (2005): Cenozoic continental climatic evolution of Central Europe. PDF file, PNAS, 102: 14964-14969. See also here.

! C. Müller et al. (2023): An integrated leaf trait analysis of two Paleogene leaf floras. In PDF, PeerJ 11: e15140 https://doi.org/10.7717/peerj.15140.
See also here.
Note figure 1: Schematic overview of the datasets used and their selection process.
Figure 6: Herbivory metrics compared between Seifhennersdorf and Suletice-Berand regarding whole assemblages and fossil-species phenology.
"... This study presents the Integrated Leaf Trait Analysis (ILTA), a workflow for the combined application of methodologies in leaf trait and insect herbivory analyses on fossil dicot leaf assemblages ..."

G.E. Mustoe et al. (2020): Neogene Tree Trunk Fossils from the Meshgin Shahr Area, Northwest Iran. In PDF, Geosciences, 10.
"... Mineralogic variations occur among different fossil trees and within a single trunk. These silica polymorphs resulted from a combination of processes: silica minerals precipitated in multiple episodes under differing geochemical conditions and the diagenetic transformation of an opaline parent material. ..."

M. Nishino et al. (2023): An exceptionally well.preserved monodominant fossil forest of Wataria from the lower Miocene of Japan. Free access, Scientific Reports, 13.

N.R. O'Brien et al. (2002): Microbial taphonomic processes in the fossilization of insects and plants in the late Eocene Florissant Formation, Colorado. Rocky Mountain Geology, 37: 1-11.
The link is to a version archived by the Internet Archive´s Wayback Machine.
See also here.

T.A. Ohsawa et al. (2016): Araucarian leaves and cone scales from the Loreto Formation of Río de Las Minas, Magellan Region, Chile. In PDF, Botany, 94: 805–815. See also here.

D.A. Oliva et al. (2022): First record of plant macrofossil from the Boa Vista Formation, Takutu Basin, Roraima State, Brazil. In PDF, Revista Brasileira de Paleontologia, 25: 303–321.
See also here.
"... X-ray diffractometry (XRD) and Laser induced-breakdown spectroscopy (LIBS) analysis were performed ..."

K.P. Pigg and M.L. DeVore (2010): Floristic composition and variation in late Paleocene to early Eocene floras in North America. Bulletin of Geosciences, 85: 135-154.

W.E. Piller et al. (2000): Palaeontological highlights of Austria. PDF file, Mitt. Österr. Geol. Ges., 92.

M.W. Rasser et al. (2013): The Randeck Maar: Palaeoenvironment and habitat differentiation of a Miocene lacustrine system. In PDF, Palaeogeography, Palaeoclimatology, Palaeoecology, 392: 426–453. See also here.

T. Reichgelt et al. (2022): Plant Proxy Evidence for High Rainfall and Productivity in the Eocene of Australia. Abstract, Paleoceanography and Paleoclimatology, 37. See also:
Palms at the Poles: Fossil Plants Reveal Lush Southern Hemisphere Forests in Ancient Hothouse Climate (by E. Hancock, UConn Communications, May 31, 2022).
Ancient Plants Provide Clues About Life on Earth in a Warmer Climate (by A. Smith, AZO Cleanteach, June 01, 2022):

Bill Rember, Jesse Garnett White and Susan Butts, Tertiary Research Center, Department of Geological Sciences University of Idaho, Moscow, ID: The Clarkia Flora of northern Idaho. Fossil plants from the Miocene.
Still available through the Internet Archive´s Wayback Machine.

J. Sakala (2004): The "Whole-Plant" concept in palaeobotany with examples from the Tertiary of northwestern Bohemia, Czech Republic with particular reference to fossil wood. PDF file (12.8 MB), Doctoral Thesis.
This expired link is still available through the Internet Archive´s Wayback Machine.
See also here. Further papers included:
Starting on PDF page 17: J. Sakala (2003): Podocarpoxylon helmstedtianum GOTTWALD from Kuklin (Late Eocene, Czech Republic) reinterpreted as Tetraclinoxylon vulcanense PRIVÉ Feddes Repertorium, 114: 25-29.
Starting on PDF page 25: J. Sakala and Catherine Privé-Gill(2004): Oligocene angiosperm woods from Northwestern Bohemia, Czech Republic. IAWA Journal, 25: 369-380.
Starting on PDF page 56: Z. Kvacek and J. Sakala (1999): Twig with attached leaves, fruits and seeds of Decodon (Lythraceae) from the Lower Miocene of northern Bohemia, and implications for the identification of detached leaves and seeds. Review of Palaeobotany and Palynology, 107: 201-222.

! C.S. Shi et al. (2013): Characterization of the stem anatomy of the Eocene fern Dennstaedtiopsis aerenchymata (Dennstaedtiaceae) by use of confocal laser scanning microscopy. Free access, American Journal of Botany, 100: 1626–1640.

E. Stiles et al. (2020): Cretaceous–Paleogene plant extinction and recovery in Patagonia. Free access, Paleobiology, 46: 445–469.

C. Strullu-Derrien et al. (2023): Insights into palaeobotany. Abstract, Botany Letters, DOI: 10.1080/23818107.2023.2200293
Note figure 1: Reconstruction of the Eocene flora from Anjou.

! B.H. Tiffney and S.R. Manchester (2001): The Use of Geological and Paleontological Evidence in Evaluating Plant Phylogeographic Hypotheses in the Northern Hemisphere Tertiary. Abstract, International Journal of Plant Sciences, 162. See also here (in PDF).

D. Uhl et al. (2021): Menatanthus mosbruggeri gen. nov. et sp. nov. – A flower with in situ pollen tetrads from the Paleocene maar lake of Menat (Puy-de-Dôme, France). Free access, Palaeobiodiversity and Palaeoenvironments, 101: 51–58.

S.W. Veatch and H.W. Meyer (2008); Start on PDF-page 12: History of paleontology at the Florissant fossil beds, Colorado. PDF file, In: Meyer, H.W., and Smith, D.M., eds., Paleontology of the Upper Eocene Florissant Formation, Colorado. The Geological Society of America, Special Paper 435: 1-18.
See also here.

B. Vento et al. (2023): Phylogenetic relationships in Nothofagus: The role of Antarctic fossil leaves. In PDF, Acta Palaeontologica Polonica, 68.

H. Wang et al. (2013): Fruits, seeds, and flowers from the Warman clay pit (middle Eocene Claiborne Group), western Tennessee, USA. In PDF, Palaeontologia Electronica. See also here.

! E.A. Wheeler and P. Baas (2022): Wood anatomy of modern and fossil Fagales in relation to phylogenetic hypotheses, familial classification, and patterns of character evolution. Free access, International Journal of Plant Sciences, 183.

V. Wilde and W. Riegel (2022): A middle Eocene treefall pit and its filling: a microenvironmental study from the onset of a forest mire in the Geiseltal (Germany). Open access, Palaeobiodiversity and Palaeoenvironments, 102: 237–251.
Note figure 10: Resin particles in palynological residue.

! E.A. Wheeler and P. Baas (1991): A Survey of the Fossil Record for Dicotiledonous Wood and its Significance for Evolutionary and Ecological Wood Anatomy. Free access, IAWA Bulletin n.s., 12: 275-332.
Note figure 1: Major ecophyletic trends of vessel element specialisation.

V. Wilde et al. (2021): A myricaceous male inflorescence with pollen in situ from the middle Eocene of Europe Open access, Palaeobiodiversity and Palaeoenvironments, 101: 873–883.

P. Wilf and R.M. Kooyman (2023): Do Southeast Asia's paleo-Antarctic trees cool the planet?
Note figure 2: Reference paleoglobes for the early Eocene (left), south polar view with part of Patagonia at the bottom and Australia at the top, and early Miocene (right), centered on Australia.
"... Many tree genera in the Malesian uplands have Southern Hemisphere origins, often supported by austral fossil records
[...] Paleo-Antarctic trees, in all likelihood, have helped cool the planet by occupying and contributing to the weathering and CO₂ consumption of uplifted terranes in Malesia over the past c. 15 Myr ..."

P. Wilf et al. (2023): The first Gondwanan Euphorbiaceae fossils reset the biogeographic history of the Macaranga-Mallotus clade. Open access, American Journal of Botany, 110: e16169.
"... The MMC [Macaranga-Mallotus clade], along with many other Gondwanan survivors, most likely entered Asia during the Neogene Sahul-Sunda collision. Our discovery adds to a substantial series of well-dated, well-preserved fossils from one undersampled region, Patagonia, that have changed our understanding of plant biogeographic history ..."
Also worth checking out:
Spurge purge: Plant fossils reveal ancient South America-to-Asia ‘escape route’. By Francisco Tutella, The Pennsylvania State University, July 24, 2023.

! P. Wilf et al. (2023): The first Gondwanan Euphorbiaceae fossils reset the biogeographic history of the Macaranga-Mallotus clade. In PDF, American Journal of Botany, e16169.

P. Wilf et al. (2019): Response to Comment on “Eocene Fagaceae from Patagonia and Gondwanan legacy in Asian rainforests”. Free access, Science. DOI: 10.1126/science.aaz2297

P. Wilf et al. (2019): Eocene Fagaceae from Patagonia and Gondwanan legacy in Asian rainforests. Abstract, Science, 364. DOI: 10.1126/science.aaw5139. See also here.

! J.A. Wolfe and G.R. Upchurch (1987): Leaf assemblages across the Cretaceous-Tertiary boundary in the Raton Basin, New Mexico and Colorado. Free access, Proc. National Academy of Sciences USA, 84: 5096-5100.

! K. Wolkenstein and G. Arp (2021): Taxon- and senescence-specific fluorescence of colored leaves from the Pliocene Willershausen Lagerstätte, Germany. Open access, PalZ.

G. Worobiec and B. Erdei (2023): The first fossil record of the anamorphic genus Zygosporium Mont. from the Oligocene of Csolnok (N Hungary). Open access, Mycological Progress, 22.

G. Worobiec and E. Worobiec 2019): Wetland vegetation from the Miocene deposits of the Belchatów Lignite Mine (central Poland). In PDF, Palaeontologia Electronica, https://doi.org/10.26879/871.

Yale Peabody Museum of Natural History, New Haven, Connecticut:
Paleobotany. Go to:
! Compendium Index of North American Paleobotany.
The Compendium Index presently covers fossil floras from North America, including Greenland, starting in the Triassic Period and extending to Pleistocene. This is a digitized version of a card-file index of approximately 20,000 images and text of descriptions of fossil plant species, maintained at Yale Peabody Museum of Natural History as a classification and identification tool.

M.C. Zamaloa et al. (2022): The first fossil record of a giant horsetail (Equisetum, Equisetaceae) is from the Miocene of Patagonia, Argentina. In PDF, Andean Geology, 49: 273-287.
See also here.

! A.E. Zanne et al. (2014): Three keys to the radiation of angiosperms into freezing environments. In PDF, Nature. Provided by the Internet Archive´s Wayback Machine.

Z. Zhou et al. (2023): Cenozoic plants from Tibet: An extraordinary decade of discovery, understanding and implications. In PDF, Science China Earth Sciences, 66: 205–226.
See also here.
Note figure 5: A reconstruction of the Eocene in the central Tibetan Plateau.